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Shinohara, "The Unfolded Protein Response in Lens Epithelial Cells from Galactosemic Rat Lenses", Invest

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In addition, ZnT5 and ZnT6 form heterodimers localized to the Golgi and ER, where they supply zinc to the early secretory pathway for metallation of enzymes such as alkaline phosphatases 33

Further, the NADH from increased glycolysis in the cytosol can be transferred to mitochondria to drive NADH-dependent NNT