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GSH can then be synthesized de novo in a two-step ATP-requiring enzymatic process utilizing glutamate cysteine ligase (GCL) and glutathione synthetase (GS) (Figure 2) [83,87]

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Four additional stabilizing mutations were added using the Quickchange multisite-directed mutagenesis kit (Agilent) to make the HexaPro variant with improved stability 71 , referred to as stabilized S trimer throughout the manuscript