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53 African populations demonstrate unique genetic advantages through higher frequencies of BCL2 protective variants promoting melanocyte survival, distinct ASIP gene polymorphisms affecting melanin production, and population-specific KIT ligand variants influencing MSCs maintenance

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73 QTL were detected to be associated with grain pericarp (exocarp/mesocarp) color, testa pigmentation, and tannin content (47 potentially novel), with key QTL colocalizing with flavonoid biosynthesis pathway genes homologous to Arabidopsis ( TT2 , TT7 , TT12 , TT16 , and AT5G41220/GST ) and rice ( MYB61 , OsbHLH025 ) (Zhang et al., 2023b)