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Zonulin was shown to reversibly open tight-junctions in a protease-activated receptor 2 (PAR2) and epidermal growth factor receptor (EGFR) dependent manner resulting in dislocation of ZO-1 from the tight junction (71)

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However, issues such as the relationship between the pathways, the existence of reciprocal regulatory roles of different pathways, and the rules of mitophagy receptor regulation under different physiological conditions remain to be resolved