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Human Embryonic Stem Cell Culture (HES) HES2 cells (ES International, Singapore) were cultured on irradiated feeders in Knock-out DMEM supplemented with 20% Knock-out Serum Replacement, 2 mmol/L glutamine, 1% non-essential amino acids (NEAA), 100 U/mL penicillin, and 100 g/mL streptomycin (all Thermo Scientific) and further supplemented with 10 ng/mL FGF2 (PeproTech)