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The phenotypes are not correlated with microenvironmental features, including proliferation rate, immune cell infiltration and vascularization, are retained when patient-derived cells are grown in vitro or as orthotopically implanted xenografts and are robust to changes in oxygen concentration, demonstrating their cell-intrinsic nature

10.1038/s41583-022-00558-9 65 de MagalhesJ

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These cells are characteristically found in hereditary spherocytosis due to cytoskeletal membrane defects, or in autoimmune hemolytic anemia where macrophages remove portions of antibody-coated erythrocyte membranes