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Cells were resuspended in 5 to 10 mL of lysis buffer (20 mM sodium phosphate pH 7.5, 200 mM NaCl, 0.05% (w/v) Tween-20, 2 mM MgCl 2 , 0.5 mM TCEP, +400 U DNAse I, 12.5 mM imidazole and a tablet of Complete EDTA-free protease inhibitor cocktail, Roche, per 100 mL of buffer) per gram of cells

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Acknowledgments The authors sincerely acknowledge the invaluable support from the Faculty of Animal Science and Technology, Yunnan Agricultural University, the Key Laboratory of Animal Nutrition and Feed Science of Yunnan Province, Yunnan Agricultural University, and the Centre for Ruminant Nutrition and Feed Engineering Technology Research, College of Animal Science and Technology, Nanjing Agricultural University