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This includes an inward-open conformation with an intact and endogenously phosphorylated R-domain tightly bound to NBD1

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Notably, alternative splicing events of a significant number of genes associated with ribosomal translation were affected by NCBP1-HNEylation, including eukaryotic translation initiation factor 4E-binding protein 1 (4ebp1 (also known as elf4ebp1))