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* Bio-GEE : biotinylated glutathione ethyl ester DTT : 1,4-dithiothreitol ETC : electron transport chain GSH : reduced glutathione GSSG : oxidized glutathione LDH : lactate dehydrogenase LPS : lipopolysaccharide MitoCDNB : Mitochondrially-targeted 1-chloro-2,4-dinitrobenzene NEM : N-ethylmaleimide OGDHC : 2-oxoglutarate dehydrogenase complex PDC : pyruvate dehydrogenase complex PDCE2 : PDC E2 component PDCE3 : PDC E3 component ROS : reactive oxygen species SDS-PAGE : sodium dodecyl sulfate-polyacrylamide gel electrophoresis TLR : toll-like receptor

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Conclusion These findings highlight the activation of native plant beneficial soil microbes as a viable strategy to mitigate co-contamination and enhance grain safety in Cd-contaminated alkaline soils